With the Allspark seemingly lost forever, Megatron resolved to abandon the planet, though the escape would demand a steep price. Meanwhile, Elita's efforts to liberate some prisoners encountered an unforeseen obstacle.

The mercenary Transformers discovered the Ark drifting aimlessly in the void. Just as unexpected allies appeared to free the captive Autobots, Megatron unveiled Project Nemesis.

The Autobots had found the Allspark, but their path was blocked by a derelict space station and its sole inhabitant. Meanwhile, the Decepticons were closing in on the Ark.

Megatron confronts a desperate struggle for survival. While the Autobots and Decepticons are united in opposition to Scorponok, Starscream plots his ascent to dominance.

Optimus found himself unexpectedly guided, shown a glimpse of a future yet to be. Meanwhile, Megatron encountered an entity who revealed the specific prize he needed to wrest from Optimus.

Above the world where the Allspark rests, Autobots and Decepticons clash in endless conflict, while Deseeus advances, driven by a thirst for vengeance against them both.